An inhibitor of mitochondrial complex II (SQR; IC50s = 0.017, 0.2, and 2 μM for bovine, rat, and A. suum nematode, respectively); inhibits QFR of A. suum (IC50 = 0.36 μM); selective for complex II over complexes I and III in rats and cattle and complex I in nematodes (IC50s >100 μM); has antibacterial and antifungal activity (EC50s = 35.9, 42.2, 60.4, and 50.2 μg/ml for R. solani, S. rolfsii, M. phaseolina, and F. oxysporum, respectively).