A cAMP phosphodiesterase inhibitor (IC50 = 41 μM); inhibits DNA topoisomerase I at a concentration of 45 μM in B16F10 melanoma cells; has antifungal activity against T. mentagrophyes with a MIC 1.8 μM; cytotoxic against A427 human lung and B16F10 mouse melanoma cells at concentrations of 25 and 200 μM, respectively; inhibits lung metastasis in mice